Spectacled Flying Fox
These large fruit bats from Australia and New Guinea sometimes produce tongue-clicks between 10-20 kHz. Most fruit bats rely entirely on vision and smell. The spectacled flying fox represents an unusual case where some echolocation appears in fruit bats. They navigate dense rainforest canopies primarily using excellent night vision. Their tongue-clicking behavior occurs mainly when flying through particularly dark areas. Research suggests this behavior varies between individuals and populations.